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Pollen flow in fragmented populations of Myrtus communis and Pistacia lentiscus: the importance of mating and pollination systems and the landscape context

Sofia Vilaça Nora, University of Seville. Supervisors: A. Aparicio and RG. Albaladejo. January 2019.
Summary:
Important questions to understand the impacts of population fragmentation are related to the genetic connectivity of populations and the key influence of pollen (and seed) dispersal of different species. Some studies have shown that the spatial distribution of populations in fragmented landscapes does not preclude incoming pollen flow even in small and isolated patches, but the low number of effective pollen sources may lead to reduced fitness in progenies. Moreover, breeding (self-compatible vs. incompatible) and pollination (wind vs. animal-pollination) systems are regarded to be very influential as well for the levels of species’ pollen flow. The thesis study system consists in populations of two common shrub species with contrasting breeding and pollination systems (Pistacia lentiscus and Myrtus communis) co-occuring in a diverse regional mosaic of mediterranean forest patches in the Guadalquivir river valley (where continuous vs. fragmented and connected vs. isolated populations can be found).

By assessing naturally produced progenies of both species in contrasting conditions of local neighbourhood conspecifics and landscape context this thesis aims to evaluate reproductive female success by performing path analysis and compare contemporary patterns of pollen flow and mating systems of Pistacia lentiscus and Myrtus communis by using indirect analysis (Twogener and Kindist), to illustrate the pollen cloud structure within and between mothers; estimate the effective distances of pollen dispersal, the effective number of pollinators and mating systems rates (selfing, outcrossing and biparental inbreeding). It also attempts to disentangle the influence of local landscapes attributes, the spatial configuration of populations itself, the blooming synchrony and the reproductive effort on the pollen flow parameters, through GLM and simulation modelling. Finally, aims to assess the importance of the pollen cloud diversity in offspring performance of both species, analysing correlations between pollen diversity and measures of fitness and survival of seedlings both under field and greenhouse conditions.