/* ============ Base ============ */
*, *::before, *::after { box-sizing: border-box; }
html { -webkit-text-size-adjust: 100%; scroll-behavior: smooth; scroll-padding-top: 112px; }
body {
  margin: 0;
  background: var(--bg);
  color: var(--text);
  font: 400 1.0625rem/1.55 var(--font-body);
  overflow-x: hidden;
  transition: background-color .3s, color .3s;
}
img { max-width: 100%; height: auto; display: block; }
a { color: inherit; }
:where(h1, h2, h3, h4) {
  font-family: var(--font-display);
  color: var(--heading);
  line-height: 1.08;
  margin: 0 0 .5em;
  text-wrap: balance;
  letter-spacing: -.02em;
}
h1 { font-weight: 800; font-size: clamp(2.5rem, 7vw, 5.5rem); }
h2 { font-weight: 800; font-size: clamp(2rem, 4.5vw, 3.25rem); }
h3 { font-weight: 600; font-size: clamp(1.2rem, 2vw, 1.5rem); }
p { margin: 0 0 1em; text-wrap: pretty; }
:focus-visible { outline: 3px solid var(--accent); outline-offset: 3px; border-radius: 4px; }
::selection { background: var(--base-a); color: #fff; }
.mono { font-family: var(--font-mono); }
.visually-hidden { position: absolute !important; width: 1px; height: 1px; overflow: hidden; clip: rect(0 0 0 0); white-space: nowrap; }
.skip-link { position: absolute; left: 16px; top: -60px; z-index: 100; background: var(--accent); color: #fff; padding: 10px 16px; border-radius: 8px; text-decoration: none; }
.skip-link:focus { top: 16px; }

.container { width: 100%; max-width: var(--maxw); margin: 0 auto; padding: 0 var(--gutter); }

/* Etiqueta mono: mayúsculas espaciadas (única excepción a "nunca todo en mayúsculas") */
.label {
  font: 500 .78rem/1 var(--font-mono);
  letter-spacing: .14em;
  text-transform: uppercase;
  color: var(--text-soft);
  display: inline-flex; align-items: center; gap: .6em;
}
.label::before { content: ""; width: 22px; height: 2px; background: currentColor; }
.label--light { color: var(--azul-300); }

/* Doble hebra: dos líneas paralelas, la inferior más corta */
.strand { display: grid; gap: 5px; width: 120px; margin: 1.25rem 0 1.75rem; }
.strand::before, .strand::after { content: ""; height: 3px; border-radius: 3px; background: var(--genoma); }
.strand::after { width: 62%; background: var(--base-a); }
.center .strand { margin-inline: auto; }

/* Tira de secuencia: las 4 primeras bases con los colores de datos */
.seq { font: 500 .85rem/1 var(--font-mono); letter-spacing: .3em; color: var(--azul-500); white-space: nowrap; }
.seq b { font-weight: 500; }
.seq .a { color: var(--base-a); } .seq .t { color: var(--base-t); }
.seq .g { color: var(--base-g); } .seq .c { color: var(--base-c); }

/* ============ Botones ============ */
.btn {
  --btn-bg: var(--genoma); --btn-fg: #fff;
  display: inline-flex; align-items: center; gap: .55em;
  padding: .85em 1.5em; border-radius: 999px; border: 2px solid transparent;
  background: var(--btn-bg); color: var(--btn-fg);
  font: 700 1rem/1 var(--font-body); text-decoration: none; cursor: pointer;
  transition: transform .25s var(--ease), box-shadow .25s, background .25s;
}
.btn:hover { transform: translateY(-2px); box-shadow: 0 10px 24px rgb(29 42 58 / .25); }
.btn svg { width: 1.1em; height: 1.1em; transition: transform .25s var(--ease); }
.btn:hover svg { transform: translateX(3px); }
.btn--accent { --btn-bg: var(--base-a-text); }
.btn--ghost { --btn-bg: transparent; --btn-fg: currentColor; border-color: currentColor; }
.btn--light { --btn-bg: #fff; --btn-fg: var(--nucleo); }

/* ============ Cabecera ============ */
.site-header {
  position: fixed; inset: 0 0 auto; z-index: 50;
  transition: background .3s, box-shadow .3s, backdrop-filter .3s;
  color: #fff;
}
.site-header.is-solid {
  background: color-mix(in srgb, var(--bg) 92%, transparent);
  backdrop-filter: blur(14px) saturate(1.4);
  -webkit-backdrop-filter: blur(14px) saturate(1.4);
  box-shadow: 0 1px 0 var(--line);
  color: var(--heading);
}
.nav { display: flex; align-items: center; gap: 1rem; height: 104px; transition: height .3s var(--ease); }
.site-header.is-solid .nav { height: 96px; }
.brand { display: flex; align-items: center; gap: .75rem; text-decoration: none; margin-right: auto; }
.brand img { height: 90px; width: auto; } /* manual: mínimo 180 px de ancho en pantalla */
.brand .logo-dark { display: none; }
.site-header.is-solid .brand .logo-light { display: none; }
.site-header.is-solid .brand .logo-dark { display: block; }
:root[data-theme="dark"] .site-header.is-solid .brand .logo-light { display: block; }
:root[data-theme="dark"] .site-header.is-solid .brand .logo-dark { display: none; }
@media (prefers-color-scheme: dark) {
  :root:not([data-theme="light"]) .site-header.is-solid .brand .logo-light { display: block; }
  :root:not([data-theme="light"]) .site-header.is-solid .brand .logo-dark { display: none; }
}
.nav-links { display: flex; gap: .25rem; list-style: none; margin: 0; padding: 0; }
.nav-links a {
  display: block; padding: .55rem .8rem; border-radius: 999px;
  text-decoration: none; font-weight: 700; font-size: .95rem; opacity: .85;
  transition: opacity .2s, background .2s;
}
.nav-links a:hover { opacity: 1; background: color-mix(in srgb, currentColor 10%, transparent); }
.nav-links a[aria-current="page"] { opacity: 1; background: color-mix(in srgb, currentColor 14%, transparent); }
.nav-tools { display: flex; gap: .4rem; align-items: center; }
.icon-btn {
  display: inline-grid; place-items: center; min-width: 40px; height: 40px; padding: 0 .6rem;
  border-radius: 999px; border: 1.5px solid color-mix(in srgb, currentColor 30%, transparent);
  background: transparent; color: inherit; cursor: pointer; font: 600 .8rem/1 var(--font-mono);
  transition: background .2s, border-color .2s;
}
.icon-btn:hover { background: color-mix(in srgb, currentColor 12%, transparent); }
.icon-btn svg { width: 18px; height: 18px; }
.menu-btn { display: none; }

@media (max-width: 960px) {
  .menu-btn { display: inline-grid; }
  .nav, .site-header.is-solid .nav { height: 80px; }
  .brand img { height: 64px; }
  .nav-links {
    position: fixed; inset: 80px 0 auto; flex-direction: column; gap: 0;
    padding: 1rem var(--gutter) 1.5rem; background: var(--bg); color: var(--heading);
    box-shadow: var(--shadow-lg); border-top: 1px solid var(--line);
    transform: translateY(-8px); opacity: 0; visibility: hidden;
    transition: opacity .25s, transform .25s var(--ease), visibility .25s;
  }
  .nav-links.is-open { transform: none; opacity: 1; visibility: visible; }
  .nav-links a { padding: .9rem .5rem; font-size: 1.15rem; border-radius: 10px; }
}

/* ============ Hero (portada) ============ */
.hero {
  position: relative; min-height: 100svh; display: grid; align-items: center;
  background:
    radial-gradient(1200px 600px at 85% 20%, rgb(70 96 127 / .45), transparent 60%),
    radial-gradient(800px 500px at 10% 90%, rgb(224 103 79 / .14), transparent 60%),
    var(--hero-bg);
  color: #fff; overflow: hidden; isolation: isolate;
}
.hero::before { /* retícula sutil de cuaderno de laboratorio */
  content: ""; position: absolute; inset: 0; z-index: -1;
  background-image: linear-gradient(rgb(255 255 255 / .04) 1px, transparent 1px), linear-gradient(90deg, rgb(255 255 255 / .04) 1px, transparent 1px);
  background-size: 48px 48px;
  mask-image: radial-gradient(ellipse at center, #000 30%, transparent 80%);
}
#helix { position: absolute; inset: 0; width: 100%; height: 100%; z-index: -1; }
.hero-content { padding-block: 140px 150px; max-width: 700px; }
.hero h1 { color: #fff; margin: 1.25rem 0 1.25rem; font-size: clamp(2.75rem, 8.5vw, 6.5rem); line-height: .95; }
.hero h1 .accent { color: var(--base-a); }
.hero-lead { font-size: clamp(1.1rem, 1.8vw, 1.35rem); color: var(--azul-200); max-width: 40ch; margin-bottom: 2rem; }
.hero-actions { display: flex; flex-wrap: wrap; gap: .75rem; }
.hero-hint { margin-top: 2.5rem; font: 500 .75rem/1.4 var(--font-mono); letter-spacing: .12em; text-transform: uppercase; color: var(--azul-300); display: flex; align-items: center; gap: .6rem; }
.hero-hint .dot { width: 8px; height: 8px; border-radius: 50%; background: var(--base-a); box-shadow: 0 0 0 0 rgb(224 103 79 / .7); animation: pulse 2s infinite; }
@keyframes pulse { 70% { box-shadow: 0 0 0 12px rgb(224 103 79 / 0); } 100% { box-shadow: 0 0 0 0 rgb(224 103 79 / 0); } }

.marquee { position: absolute; inset: auto 0 0; padding: 1rem 0; border-top: 1px solid rgb(255 255 255 / .08); overflow: hidden; }
.marquee-track { display: flex; width: max-content; animation: marquee 60s linear infinite; }
.marquee .seq { color: rgb(159 182 211 / .55); padding-right: 2rem; }
@keyframes marquee { to { transform: translateX(-50%); } }

/* Cabecera de páginas interiores */
.page-hero {
  position: relative; background: var(--hero-bg); color: #fff; overflow: hidden; isolation: isolate;
  padding: 150px 0 72px;
}
.page-hero::after {
  content: attr(data-letter); position: absolute; right: -2vw; bottom: -.18em; z-index: -1;
  font: 800 clamp(10rem, 30vw, 24rem)/1 var(--font-display); color: rgb(255 255 255 / .04);
}
.page-hero h1 { color: #fff; font-size: clamp(2.5rem, 6.5vw, 4.75rem); margin: 1rem 0 .75rem; }
.page-hero p { color: var(--azul-200); font-size: 1.2rem; max-width: 52ch; margin: 0; }
.page-hero .strand::before { background: var(--azul-300); }

/* ============ Secciones ============ */
.section { padding: clamp(72px, 10vw, 128px) 0; position: relative; }
.section--alt { background: var(--bg-alt); }
.section--dark { background: var(--nucleo); color: var(--azul-200); }
.section--dark :where(h2, h3) { color: #fff; }
.section-head { max-width: 720px; margin-bottom: clamp(2rem, 4vw, 3.5rem); }
.section-head.center { text-align: center; margin-inline: auto; }
.section-head p { font-size: 1.15rem; color: var(--text-soft); }
.section--dark .section-head p { color: var(--azul-300); }
.section-foot { margin-top: 2.5rem; display: flex; justify-content: center; }

.grid { display: grid; gap: clamp(16px, 2.5vw, 28px); }
.grid-2 { grid-template-columns: repeat(auto-fit, minmax(min(100%, 420px), 1fr)); }
.grid-3 { grid-template-columns: repeat(auto-fill, minmax(min(100%, 300px), 1fr)); }
.grid-4 { grid-template-columns: repeat(auto-fill, minmax(min(100%, 240px), 1fr)); }

/* Sobre nosotros */
.about { display: grid; grid-template-columns: 1.05fr 1fr; gap: clamp(32px, 6vw, 80px); align-items: center; }
.about-text p { font-size: 1.15rem; }
.about-text strong { color: var(--heading); }
.about-photo { position: relative; }
.about-photo img { border-radius: var(--radius-lg); box-shadow: var(--shadow-lg); aspect-ratio: 16 / 10; object-fit: cover; width: 100%; height: auto; }
.about-photo figcaption { margin-top: .75rem; font: .8rem/1.4 var(--font-mono); color: var(--text-soft); }
.about-photo::before {
  content: ""; position: absolute; inset: -18px 18px 18px -18px; z-index: -1;
  border: 2px solid var(--azul-200); border-radius: var(--radius-lg);
}
@media (max-width: 860px) { .about { grid-template-columns: 1fr; } }

/* Contadores */
.stats { display: grid; grid-template-columns: repeat(4, 1fr); gap: 1px; background: var(--line); border-radius: var(--radius-lg); overflow: hidden; margin-top: clamp(48px, 7vw, 88px); border: 1px solid var(--line); }
.stat { background: var(--surface); padding: 1.75rem 1.5rem; }
.stat-num { font: 800 clamp(2.5rem, 5vw, 3.5rem)/1 var(--font-display); color: var(--heading); display: block; }
.stat-num sup { color: var(--base-a); font-size: .5em; }
.stat-label { font: 500 .75rem/1.3 var(--font-mono); letter-spacing: .12em; text-transform: uppercase; color: var(--text-soft); margin-top: .6rem; display: block; }
@media (max-width: 720px) { .stats { grid-template-columns: repeat(2, 1fr); } }

/* Líneas de investigación */
.line-card {
  --c: var(--genoma); --c-text: var(--genoma);
  position: relative; display: flex; flex-direction: column; gap: .75rem;
  padding: 2rem 1.75rem 1.75rem; border-radius: var(--radius-lg);
  background: var(--surface); border: 1px solid var(--line); box-shadow: var(--shadow);
  text-decoration: none; overflow: hidden; isolation: isolate;
  transition: transform .4s var(--ease), box-shadow .4s, border-color .4s;
}
.line-card::before {
  content: ""; position: absolute; inset: 0; z-index: -1; opacity: 0;
  background: radial-gradient(400px 220px at 100% 0%, color-mix(in srgb, var(--c) 22%, transparent), transparent 70%);
  transition: opacity .4s;
}
.line-card:hover { transform: translateY(-6px); box-shadow: var(--shadow-lg); border-color: color-mix(in srgb, var(--c) 50%, var(--line)); }
.line-card:hover::before { opacity: 1; }
.line-base {
  width: 56px; height: 56px; border-radius: 16px; display: grid; place-items: center;
  background: var(--c); color: #fff; font: 800 1.75rem/1 var(--font-display);
  transition: transform .5s var(--ease);
}
.line-card:hover .line-base { transform: rotate(-8deg) scale(1.08); }
.line-card .label { color: var(--c-text); }
.line-card h3 { margin: 0; }
.line-card p { color: var(--text-soft); margin: 0; }
.line-card ul { margin: .25rem 0 0; padding: 0; list-style: none; display: grid; gap: .4rem; }
.line-card li { display: flex; gap: .6rem; font-size: .97rem; }
.line-card li::before { content: ""; flex: none; width: 7px; height: 7px; margin-top: .55em; border-radius: 2px; background: var(--c); }
.line-more { margin-top: auto; padding-top: 1rem; font-weight: 700; color: var(--c-text); display: inline-flex; gap: .4rem; align-items: center; }
.line-more svg { width: 1.1em; height: 1.1em; transition: transform .3s var(--ease); }
.line-card:hover .line-more svg { transform: translateX(4px); }
.line-focus { display: flex; flex-wrap: wrap; gap: .4rem; }
.line-focus span {
  padding: .3rem .65rem; border-radius: 999px; font: 600 .74rem/1.2 var(--font-mono); letter-spacing: .06em; text-transform: uppercase;
  color: var(--c-text); background: color-mix(in srgb, var(--c) 14%, transparent); border: 1px solid color-mix(in srgb, var(--c) 45%, transparent);
}
.research-line .line-focus { margin: .25rem 0 .5rem; }
.base-a { --c: var(--base-a); --c-text: var(--base-a-text); }
.base-t { --c: var(--base-t); --c-text: var(--base-t-text); }
.base-g { --c: var(--base-g); --c-text: var(--base-g-text); }
.base-c { --c: var(--base-c); --c-text: var(--base-c-text); }
:root[data-theme="dark"] .line-card, :root[data-theme="dark"] .research-line { --c-text: color-mix(in srgb, var(--c) 75%, #fff); }
@media (prefers-color-scheme: dark) { :root:not([data-theme="light"]) :is(.line-card, .research-line) { --c-text: color-mix(in srgb, var(--c) 75%, #fff); } }

/* Página de investigación: bloques detallados */
.research-line {
  display: grid; grid-template-columns: minmax(0, 1fr) minmax(0, 1.1fr); gap: clamp(28px, 5vw, 72px); align-items: center;
  padding: clamp(48px, 7vw, 96px) 0; border-bottom: 1px solid var(--line);
}
.research-line:last-child { border-bottom: 0; }
.research-line:nth-child(even) .research-visual { order: -1; }
.research-line .label { color: var(--c-text); }
.research-line h2 { font-size: clamp(1.9rem, 3.8vw, 2.75rem); }
.research-line .strand::after { background: var(--c); }
.research-line ul { padding: 0; list-style: none; display: grid; gap: .65rem; margin: 1.25rem 0; }
.research-line li { padding: .9rem 1.1rem; border-radius: 12px; background: var(--bg-alt); border-left: 4px solid var(--c); }
.research-visual { border-radius: var(--radius-lg); background: var(--bg-alt); border: 1px solid var(--line); padding: clamp(16px, 3vw, 32px); }
.research-visual svg { width: 100%; height: auto; display: block; }
.research-team { display: flex; flex-wrap: wrap; gap: .5rem; margin-top: 1rem; }
.chip { display: inline-flex; align-items: center; gap: .45rem; padding: .35rem .8rem .35rem .35rem; border-radius: 999px; background: var(--surface); border: 1px solid var(--line); font-size: .9rem; text-decoration: none; }
.chip img, .chip .avatar { width: 26px; height: 26px; border-radius: 50%; object-fit: cover; font-size: .65rem; }
@media (max-width: 860px) {
  .research-line { grid-template-columns: 1fr; }
  .research-line:nth-child(even) .research-visual { order: 0; }
}

/* Cita */
.quote { text-align: center; max-width: 960px; margin: 0 auto; }
.quote blockquote { margin: 0; font: 800 clamp(1.75rem, 4.5vw, 3.25rem)/1.12 var(--font-display); color: #fff; letter-spacing: -.02em; text-wrap: balance; }
.quote blockquote .hl { color: var(--base-a); }
.quote figcaption { margin-top: 1.75rem; font: 500 .85rem/1.4 var(--font-mono); letter-spacing: .12em; text-transform: uppercase; color: var(--azul-300); }
.quote-mark { font: 800 7rem/0.6 var(--font-display); color: var(--base-a); display: block; height: .55em; }

/* Publicaciones */
.pub {
  display: grid; grid-template-columns: 72px 1fr; gap: 1.25rem;
  padding: 1.5rem 0; border-top: 1px solid var(--line);
}
.pub:last-child { border-bottom: 1px solid var(--line); }
.pub-year { font: 800 1.5rem/1 var(--font-display); color: var(--azul-500); padding-top: .2rem; }
.pub-title { font: 600 1.15rem/1.3 var(--font-display); color: var(--heading); margin: 0 0 .4rem; }
.pub-title a { text-decoration: none; background: linear-gradient(var(--accent), var(--accent)) 0 100% / 0 2px no-repeat; transition: background-size .35s var(--ease); }
.pub-title a:hover { background-size: 100% 2px; }
.pub-authors { font-size: .95rem; color: var(--text-soft); margin: 0 0 .5rem; }
.pub-authors strong { color: var(--heading); }
.pub-meta { display: flex; flex-wrap: wrap; gap: .5rem; align-items: center; font: .8rem/1 var(--font-mono); }
.tag { display: inline-flex; align-items: center; gap: .35rem; padding: .35rem .6rem; border-radius: 6px; background: var(--bg-alt); color: var(--text-soft); text-decoration: none; border: 1px solid var(--line); }
a.tag:hover { border-color: var(--genoma); color: var(--heading); }
.tag--journal { background: var(--genoma); color: #fff; border-color: transparent; }
.tag--key { background: var(--base-a-tint); color: var(--base-a-text); border-color: transparent; }
@media (max-width: 560px) { .pub { grid-template-columns: 1fr; gap: .5rem; } }

.pub-card { display: flex; flex-direction: column; gap: .75rem; padding: 1.75rem; border-radius: var(--radius-lg); background: var(--surface); border: 1px solid var(--line); box-shadow: var(--shadow); transition: transform .35s var(--ease), box-shadow .35s; }
.pub-card:hover { transform: translateY(-4px); box-shadow: var(--shadow-lg); }
.pub-card .pub-title { font-size: 1.2rem; }
.pub-card .pub-meta { margin-top: auto; }

.toolbar { display: flex; flex-wrap: wrap; gap: .75rem; align-items: center; margin-bottom: 2rem; }
.search {
  flex: 1 1 260px; display: flex; align-items: center; gap: .6rem; padding: 0 1rem; height: 48px;
  border-radius: 999px; border: 1.5px solid var(--line); background: var(--surface);
}
.search:focus-within { border-color: var(--genoma); }
.search input { flex: 1; border: 0; background: transparent; color: var(--text); font: inherit; outline: none; min-width: 0; }
.search svg { width: 18px; height: 18px; color: var(--text-soft); flex: none; }
.chips { display: flex; flex-wrap: wrap; gap: .4rem; }
.chip-btn {
  padding: .55rem 1rem; border-radius: 999px; border: 1.5px solid var(--line); background: var(--surface);
  color: var(--text); font: 600 .9rem/1 var(--font-body); cursor: pointer; transition: all .2s;
}
.chip-btn:hover { border-color: var(--genoma); }
.chip-btn[aria-pressed="true"] { background: var(--nucleo); border-color: var(--nucleo); color: #fff; }
:root[data-theme="dark"] .chip-btn[aria-pressed="true"] { background: var(--azul-200); border-color: var(--azul-200); color: var(--nucleo); }
@media (prefers-color-scheme: dark) { :root:not([data-theme="light"]) .chip-btn[aria-pressed="true"] { background: var(--azul-200); border-color: var(--azul-200); color: var(--nucleo); } }
.empty { padding: 3rem 1rem; text-align: center; color: var(--text-soft); border: 1.5px dashed var(--line); border-radius: var(--radius-lg); }
.result-count { font: .8rem/1 var(--font-mono); color: var(--text-soft); letter-spacing: .08em; }

/* Equipo */
.team-group + .team-group { margin-top: clamp(48px, 6vw, 72px); }
.team-group > h2 { font-size: clamp(1.5rem, 3vw, 2rem); display: flex; align-items: baseline; gap: .75rem; }
.team-group > h2 .count { font: 500 .85rem/1 var(--font-mono); color: var(--text-soft); }
.member { position: relative; border-radius: var(--radius-lg); overflow: hidden; background: var(--surface); border: 1px solid var(--line); box-shadow: var(--shadow); transition: transform .4s var(--ease), box-shadow .4s; }
.member:hover { transform: translateY(-6px); box-shadow: var(--shadow-lg); }
.member-photo { position: relative; aspect-ratio: 4 / 5; overflow: hidden; background: var(--bg-alt); }
.member-photo img { width: 100%; height: 100%; object-fit: cover; transition: transform .7s var(--ease); }
.member:hover .member-photo img { transform: scale(1.05); }
.member-photo::after { /* tira ATGC al pasar el ratón */
  content: ""; position: absolute; inset: auto 0 0; height: 5px;
  background: linear-gradient(90deg, var(--base-a) 0 25%, var(--base-t) 0 50%, var(--base-g) 0 75%, var(--base-c) 0);
  transform: scaleX(0); transform-origin: left; transition: transform .5s var(--ease);
}
.member:hover .member-photo::after { transform: scaleX(1); }
.member-body { padding: 1.1rem 1.25rem 1.3rem; }
.member-name { font: 600 1.15rem/1.2 var(--font-display); color: var(--heading); margin: 0 0 .3rem; }
.member-role { font-size: .92rem; color: var(--text-soft); margin: 0; }
.member-links { display: flex; gap: .4rem; margin-top: .75rem; }
.member-links a { font: .72rem/1 var(--font-mono); padding: .35rem .55rem; border-radius: 6px; background: var(--bg-alt); text-decoration: none; color: var(--text-soft); }
.member-links a:hover { color: var(--heading); }
.member--pi { grid-column: 1 / -1; max-width: 920px; display: grid; grid-template-columns: minmax(220px, 340px) minmax(0, 1fr); }
.member--pi .member-photo { aspect-ratio: auto; min-height: 100%; }
.member--pi .member-body { padding: clamp(1.5rem, 3vw, 2.5rem); display: flex; flex-direction: column; justify-content: center; }
.member--pi .member-name { font-size: clamp(1.6rem, 3vw, 2.2rem); font-weight: 800; }
.member--pi .member-bio { margin-top: 1rem; color: var(--text); font-size: 1.1rem; max-width: 52ch; }
@media (max-width: 640px) { .member--pi { grid-column: auto; grid-template-columns: 1fr; } .member--pi .member-photo { aspect-ratio: 4 / 5; } }
.avatar {
  width: 100%; height: 100%; display: grid; place-items: center;
  font: 800 clamp(2.5rem, 7vw, 4rem)/1 var(--font-display); color: #fff; letter-spacing: -.03em;
  background: var(--genoma);
}
.avatar[data-base="a"] { background: linear-gradient(135deg, var(--genoma), var(--base-a-text)); }
.avatar[data-base="t"] { background: linear-gradient(135deg, var(--genoma), var(--base-t-text)); }
.avatar[data-base="g"] { background: linear-gradient(135deg, var(--nucleo), var(--base-g-text)); }
.avatar[data-base="c"] { background: linear-gradient(135deg, var(--nucleo), var(--base-c-text)); }

/* Galería */
.gallery { columns: 3 280px; column-gap: 16px; }
.gallery button { all: unset; display: block; margin-bottom: 16px; border-radius: var(--radius); overflow: hidden; cursor: zoom-in; position: relative; break-inside: avoid; }
.gallery button:focus-visible { outline: 3px solid var(--accent); outline-offset: 3px; }
.gallery img { width: 100%; height: auto; transition: transform .6s var(--ease); }
.gallery button:hover img { transform: scale(1.04); }
.gallery figcaption { position: absolute; inset: auto 0 0; padding: 2rem 1rem .8rem; color: #fff; font-size: .9rem; background: linear-gradient(transparent, rgb(29 42 58 / .85)); opacity: 0; transition: opacity .3s; }
.gallery button:hover figcaption, .gallery button:focus-visible figcaption { opacity: 1; }
.lightbox { position: fixed; inset: 0; z-index: 200; display: grid; place-items: center; padding: 24px; background: rgb(15 23 34 / .92); opacity: 0; visibility: hidden; transition: opacity .3s, visibility .3s; }
.lightbox.is-open { opacity: 1; visibility: visible; }
.lightbox img { max-height: 82vh; max-width: min(1200px, 100%); width: auto; border-radius: var(--radius); box-shadow: var(--shadow-lg); }
.lightbox p { color: var(--azul-200); text-align: center; margin: 1rem 0 0; }
.lightbox .icon-btn { position: absolute; color: #fff; background: rgb(255 255 255 / .08); }
.lb-close { top: 20px; right: 20px; }
.lb-prev { left: 16px; top: 50%; } .lb-next { right: 16px; top: 50%; }

/* Noticias */
.news-card { display: flex; flex-direction: column; border-radius: var(--radius-lg); overflow: hidden; background: var(--surface); border: 1px solid var(--line); box-shadow: var(--shadow); transition: transform .35s var(--ease), box-shadow .35s; }
.news-card:hover { transform: translateY(-4px); box-shadow: var(--shadow-lg); }
.news-media { aspect-ratio: 16 / 9; background: var(--nucleo); overflow: hidden; position: relative; }
.news-media img { width: 100%; height: 100%; object-fit: cover; }
.news-media { background: linear-gradient(135deg, var(--nucleo-deep), var(--genoma)); }
.news-pattern { position: absolute; inset: -6px -10px; display: block; font-size: .95rem; line-height: 1.55; letter-spacing: .32em; color: rgb(201 215 234 / .22); white-space: nowrap; overflow: hidden; transition: transform 1.2s var(--ease); }
.news-pattern b { opacity: 1; }
.news-card:hover .news-pattern { transform: translateX(-24px); }
.news-body { padding: 1.4rem 1.5rem 1.6rem; display: flex; flex-direction: column; gap: .55rem; flex: 1; }
.news-body h3 { margin: 0; font-size: 1.25rem; }
.news-body p { margin: 0; color: var(--text-soft); }
.news-date { font: 500 .75rem/1 var(--font-mono); letter-spacing: .1em; text-transform: uppercase; color: var(--accent-text); }
.news-body .btn { margin-top: auto; align-self: flex-start; padding: .6em 1.1em; font-size: .9rem; }
.news-cat { font: .7rem/1 var(--font-mono); letter-spacing: .1em; text-transform: uppercase; padding: .35rem .55rem; border-radius: 6px; background: rgb(255 255 255 / .9); color: var(--nucleo); position: absolute; top: 12px; left: 12px; }

/* Llamada a la acción */
.cta {
  position: relative; overflow: hidden; isolation: isolate;
  border-radius: var(--radius-lg); padding: clamp(32px, 6vw, 72px);
  background: linear-gradient(120deg, var(--nucleo), var(--genoma)); color: #fff;
  display: grid; grid-template-columns: 1.4fr auto; gap: 2rem; align-items: center;
}
.cta::after { content: "ATGC"; position: absolute; right: -1rem; bottom: -3rem; z-index: -1; font: 800 12rem/1 var(--font-display); color: rgb(255 255 255 / .06); letter-spacing: -.04em; }
.cta h2 { color: #fff; margin-bottom: .4em; }
.cta p { color: var(--azul-200); margin: 0; font-size: 1.1rem; max-width: 54ch; }
@media (max-width: 760px) { .cta { grid-template-columns: 1fr; } }

/* Financiación */
.funders { display: flex; flex-wrap: wrap; gap: 12px; justify-content: center; }
.funder { display: inline-flex; align-items: center; justify-content: center; min-height: 72px; padding: .8rem 1.25rem; border-radius: var(--radius); background: var(--surface); border: 1px solid var(--line); text-decoration: none; font: 600 .95rem/1.25 var(--font-display); color: var(--heading); text-align: center; transition: border-color .2s, transform .3s var(--ease); }
.funder:hover { border-color: var(--genoma); transform: translateY(-2px); }
.funder img { max-height: 56px; width: auto; }

/* Contacto */
.contact-grid { display: grid; grid-template-columns: 1fr 1fr; gap: clamp(24px, 4vw, 48px); }
.contact-card { padding: clamp(24px, 4vw, 40px); border-radius: var(--radius-lg); background: var(--surface); border: 1px solid var(--line); box-shadow: var(--shadow); }
.contact-card dl { margin: 0; display: grid; gap: 1.25rem; }
.contact-card dt { font: 500 .75rem/1 var(--font-mono); letter-spacing: .12em; text-transform: uppercase; color: var(--text-soft); margin-bottom: .4rem; }
.contact-card dd { margin: 0; font-size: 1.1rem; }
.contact-card a { color: var(--accent-text); font-weight: 700; }
.job { padding: 1.5rem; border-radius: var(--radius); border: 1px solid var(--line); background: var(--bg-alt); }
.job + .job { margin-top: 1rem; }
.job h3 { margin-bottom: .4rem; }
.job ul { margin: .5rem 0 0; padding-left: 1.1rem; color: var(--text-soft); }
@media (max-width: 860px) { .contact-grid { grid-template-columns: 1fr; } }
.map-embed { margin-top: 1.75rem; aspect-ratio: 4 / 3; border-radius: var(--radius); overflow: hidden; border: 1px solid var(--line); background: linear-gradient(135deg, var(--nucleo-deep), var(--genoma)); }
.map-embed iframe { display: block; width: 100%; height: 100%; border: 0; }
.map-placeholder { height: 100%; display: grid; place-content: center; justify-items: center; gap: .9rem; padding: 1.5rem; text-align: center; color: #fff; }
.map-placeholder > svg { width: 44px; height: 44px; color: var(--azul-200); }
.contact-card .map-placeholder .btn { color: var(--nucleo); }
.map-consent { margin: 0; max-width: 30ch; font-size: .8rem; color: var(--azul-200); }

/* Divulgación: laboratorio interactivo de reparación */
.lab-game { border-radius: var(--radius-lg); background: var(--nucleo-deep); color: var(--azul-200); padding: clamp(20px, 4vw, 40px); box-shadow: var(--shadow-lg); }
.lab-game h3 { color: #fff; }
.game-stage { position: relative; border-radius: var(--radius); background: #0F1722; border: 1px solid rgb(255 255 255 / .08); overflow: hidden; }
.game-stage svg { display: block; width: 100%; height: auto; }
.game-controls { display: flex; flex-wrap: wrap; gap: .6rem; margin-top: 1.25rem; align-items: center; }
.game-controls .btn { padding: .7em 1.2em; font-size: .95rem; }
.game-controls .btn[disabled] { opacity: .4; pointer-events: none; }
.phase-toggle { display: inline-flex; border-radius: 999px; border: 1.5px solid rgb(255 255 255 / .2); padding: 3px; margin-right: auto; }
.phase-toggle button { all: unset; cursor: pointer; padding: .5rem .9rem; border-radius: 999px; font: 600 .8rem/1 var(--font-mono); color: var(--azul-300); }
.phase-toggle button[aria-pressed="true"] { background: var(--azul-200); color: var(--nucleo); }
.phase-toggle button:focus-visible { outline: 2px solid var(--base-a); }
.game-log { margin-top: 1.25rem; min-height: 4.5em; padding: 1rem 1.2rem; border-radius: var(--radius); background: rgb(255 255 255 / .05); border-left: 4px solid var(--azul-500); }
.game-log strong { color: #fff; }
.game-log.is-good { border-color: var(--base-t); }
.game-log.is-bad { border-color: var(--base-a); }
.game-score { font: .8rem/1.5 var(--font-mono); color: var(--azul-300); letter-spacing: .08em; margin: .9rem 0 0; }


/* ============ Pie ============ */
.site-footer { background: var(--nucleo-deep); color: var(--azul-300); padding: 72px 0 32px; }
.footer-grid { display: grid; grid-template-columns: 1.4fr 1fr 1fr; gap: 40px; }
.site-footer img { height: 90px; width: auto; margin-bottom: 1rem; }
.site-footer h4 { font: 500 .75rem/1 var(--font-mono); letter-spacing: .14em; text-transform: uppercase; color: #fff; margin-bottom: 1rem; }
.site-footer ul { list-style: none; padding: 0; margin: 0; display: grid; gap: .5rem; }
.site-footer a { text-decoration: none; color: var(--azul-200); }
.site-footer a:hover { color: #fff; text-decoration: underline; text-decoration-color: var(--base-a); text-underline-offset: 4px; }
.footer-bottom { margin-top: 56px; padding-top: 24px; border-top: 1px solid rgb(255 255 255 / .08); display: flex; flex-wrap: wrap; gap: 1rem; justify-content: space-between; align-items: center; font-size: .85rem; }
@media (max-width: 760px) { .footer-grid { grid-template-columns: 1fr; } }
.footer-cookies { background: none; border: 0; padding: 0; font: inherit; color: inherit; text-decoration: underline; text-underline-offset: 3px; cursor: pointer; }
.footer-cookies:hover { color: #fff; }

/* Banner de cookies */
.cookie-banner {
  position: fixed; left: 16px; right: 16px; bottom: 16px; z-index: 150; margin-inline: auto; max-width: 880px;
  display: flex; flex-wrap: wrap; gap: 1rem 1.75rem; align-items: center; justify-content: space-between;
  padding: 1.25rem 1.5rem; border-radius: var(--radius-lg); background: var(--surface); color: var(--text);
  border: 1px solid var(--line); box-shadow: var(--shadow-lg); animation: cookie-in .5s var(--ease) both;
}
.cookie-text { flex: 1 1 360px; }
.cookie-text p { margin: .5rem 0 0; font-size: .92rem; }
.cookie-actions { display: flex; gap: .75rem; flex-wrap: wrap; }
.cookie-actions .btn { min-width: 8.5rem; justify-content: center; }
@keyframes cookie-in { from { opacity: 0; transform: translateY(24px); } }
@media (prefers-reduced-motion: reduce) { .cookie-banner { animation: none; } }
@media (max-width: 520px) { .cookie-actions { width: 100%; } .cookie-actions .btn { flex: 1; min-width: 0; } }

/* ============ Animaciones de entrada ============ */
[data-reveal] { opacity: 0; transform: translateY(28px); transition: opacity .8s var(--ease), transform .8s var(--ease); transition-delay: var(--d, 0s); }
[data-reveal].is-in { opacity: 1; transform: none; }
.no-js [data-reveal] { opacity: 1; transform: none; }

@media (prefers-reduced-motion: reduce) {
  html { scroll-behavior: auto; }
  *, *::before, *::after { animation-duration: .01ms !important; animation-iteration-count: 1 !important; transition-duration: .01ms !important; }
  [data-reveal] { opacity: 1; transform: none; }
}

/* ============ Diagramas de investigación ============ */
.dg { overflow: visible; }
.dg-cell { fill: var(--surface); stroke: var(--azul-200); stroke-width: 2; }
.dg-soft { fill: color-mix(in srgb, var(--genoma) 18%, transparent); }
.dg-dna { fill: none; stroke: var(--genoma); stroke-width: 4; stroke-linecap: round; }
.dg-rna { fill: none; stroke: var(--c); stroke-width: 3; stroke-linecap: round; stroke-dasharray: 160; stroke-dashoffset: 160; animation: dg-draw 2.4s var(--ease) infinite alternate; }
.dg-bead { fill: var(--genoma); stroke: var(--surface); stroke-width: 3; }
.dg-tail { stroke: var(--azul-500); stroke-width: 2; }
.dg-mark { fill: var(--c); }
.dg-cap { fill: var(--c); }
.dg-cap-zone { fill: none; stroke: var(--azul-300); stroke-width: 2; stroke-dasharray: 6 6; }
.dg-bolt { fill: var(--c); }
.dg-pulse { fill: none; stroke: var(--c); stroke-width: 2; transform-box: fill-box; transform-origin: center; animation: dg-pulse 2s ease-out infinite; }
.dg-protein { fill: var(--nucleo); }
:root[data-theme="dark"] .dg-protein { fill: var(--azul-500); }
.dg-label-in { fill: #fff; font: 600 13px var(--font-mono); }
.dg-arrow { fill: none; stroke: var(--azul-500); stroke-width: 2; stroke-dasharray: 5 5; }
.dg-label text, .dg-label-small { fill: var(--text-soft); font: 500 11px var(--font-mono); letter-spacing: .08em; }
.dg-label line { stroke: var(--azul-300); stroke-width: 1.2; }
.dg-label .dg-key, .dg .dg-key { fill: var(--c-text); font-weight: 700; }
.dg-ray { fill: none; stroke: var(--c); stroke-width: 3; stroke-linejoin: round; }
.dg-vessel { fill: none; stroke: var(--azul-300); stroke-width: 2.5; }
.dg-vesicle { fill: var(--c); opacity: .85; animation: dg-float 2.4s ease-in-out infinite alternate; }
.dg-tube { fill: var(--surface); stroke: var(--genoma); stroke-width: 2.5; }
.dg-tube-fill { fill: color-mix(in srgb, var(--c) 55%, transparent); }
.dg-box { fill: var(--surface); stroke: var(--azul-300); stroke-width: 2; }
.dg-box--key { stroke: var(--c); stroke-width: 2.5; }
@keyframes dg-pulse { from { transform: scale(.6); opacity: 1; } to { transform: scale(2.2); opacity: 0; } }
@keyframes dg-draw { to { stroke-dashoffset: 0; } }
@keyframes dg-float { to { transform: translate(6px, -5px); } }
@media (prefers-color-scheme: dark) { :root:not([data-theme="light"]) .dg-protein { fill: var(--azul-500); } }
@media (max-width: 999px) { .hero-content { padding-bottom: 240px; } }

/* ============ Juego "Repara el ADN" ============ */
.gm-seg { stroke: var(--azul-300); stroke-width: 5; stroke-linecap: round; transition: opacity .5s; }
.gm-rung { stroke-width: 6; stroke-linecap: round; transition: opacity .5s; }
.gm-letter { fill: var(--azul-200); font: 600 13px var(--font-mono); text-anchor: middle; transition: opacity .5s; }
.is-scar .gm-letter { fill: var(--base-a); }
.is-scar .gm-seg { stroke: var(--base-a); }
.gm-half { transition: transform .6s var(--ease); transform-box: fill-box; }
.gm-left { transform-origin: right center; }
.gm-right { transform-origin: left center; }
.is-broken .gm-left { transform: translateX(-30px) rotate(-3deg); }
.is-broken .gm-right { transform: translateX(30px) rotate(3deg); }
.is-shaking .gm-half { animation: gm-shake .5s; }
.gm-glow { fill: rgb(224 103 79 / .18); opacity: 0; transition: opacity .4s; }
.is-broken .gm-glow { opacity: 1; animation: gm-glow 1.4s ease-in-out infinite alternate; }
.gm-sister { opacity: .1; transition: opacity .6s; }
.gm-sister.is-on { opacity: .55; }
.is-invading .gm-sister.is-on { opacity: 1; }
.is-invading .gm-sister .gm-seg { stroke: var(--base-t); }
.gm-caption { fill: var(--azul-300); font: 500 11px var(--font-mono); letter-spacing: .1em; }
.gm-invade { fill: none; stroke: var(--base-t); stroke-width: 3; stroke-dasharray: 6 6; opacity: 0; transition: opacity .4s; }
.is-invading .gm-invade { opacity: 1; animation: gm-dash 1s linear infinite; }
.is-resected-top .gm-top, .is-resected-bot .gm-bot { opacity: .08; }
.is-copied .gm-seg { stroke: var(--base-t); }
.is-copied .gm-letter { fill: var(--base-t); }
@keyframes gm-shake { 25% { transform: translateX(-36px) rotate(-3deg); } 75% { transform: translateX(-24px) rotate(-3deg); } }
.is-shaking .gm-right { animation: gm-shake-r .5s; }
@keyframes gm-shake-r { 25% { transform: translateX(36px) rotate(3deg); } 75% { transform: translateX(24px) rotate(3deg); } }
@keyframes gm-glow { to { r: 38; } }
@keyframes gm-dash { to { stroke-dashoffset: -24; } }
.dg-repeat { stroke-width: 5; stroke-dasharray: 14 5; }
.dg-folia { fill: none; stroke: var(--azul-300); stroke-width: 1.5; stroke-linecap: round; }
.dg-tumor { fill: var(--c); }
.dg-cc { fill: none; stroke: var(--azul-300); stroke-width: 6; stroke-linecap: round; }
.dg-lens { fill: var(--surface); stroke: var(--c); stroke-width: 2; }
.dg-mark-lost { fill: none; stroke: var(--c); stroke-width: 1.5; stroke-dasharray: 2 2; }
.dg-mark-fade { fill: var(--c); opacity: 0; animation: dg-fade 2.4s ease-in-out infinite alternate; }
@keyframes dg-fade { from { opacity: 1; } 60%, to { opacity: 0; } }
.dg-repeat-box { fill: color-mix(in srgb, var(--c) 30%, var(--surface)); stroke: var(--c); stroke-width: 2; }
.dg-read { stroke: var(--azul-300); stroke-width: 4; stroke-linecap: round; }
.dg-chev { fill: none; stroke: var(--c-text); stroke-width: 2; stroke-linecap: round; stroke-linejoin: round; }
.dg-aa { fill: var(--c); transform-box: fill-box; transform-origin: center; animation: dg-bob 1.6s ease-in-out infinite alternate; }
@keyframes dg-bob { to { transform: translateY(-4px); } }

/* ============ Enlaces con icono (personas, redes) ============ */
.member-links { flex-wrap: wrap; }
.member-links .link-btn, .footer-social .link-btn {
  display: inline-grid; place-items: center; width: 36px; height: 36px; padding: 0; border-radius: 10px;
  background: var(--bg-alt); color: var(--text-soft); border: 1px solid var(--line);
  transition: transform .25s var(--ease), color .2s, background .2s, border-color .2s;
}
.link-btn svg { width: 17px; height: 17px; }
.link-btn:hover { transform: translateY(-2px); color: #fff; }
.link-btn--linkedin:hover { background: #0A66C2; border-color: #0A66C2; }
.link-btn--x:hover { background: #000; border-color: #000; }
.link-btn--instagram:hover { background: linear-gradient(45deg, #F58529, #DD2A7B 55%, #8134AF); border-color: transparent; }
.link-btn--email:hover, .link-btn--web:hover, .link-btn--scholar:hover, .link-btn--youtube:hover { background: var(--genoma); border-color: var(--genoma); }
.member-links .link-btn--pubmed, .footer-social .link-btn--pubmed { width: auto; padding: 0 .7rem; }
.link-btn--pubmed:hover { background: #fff; border-color: #20558A; }

/* Logotipo tipográfico de PubMed */
.pubmed-mark { font: 700 .82rem/1 Arial, "Helvetica Neue", sans-serif; letter-spacing: -.01em; color: #20558A; }
.pubmed-mark b { color: #4D8DCA; font-weight: 700; }
:root[data-theme="dark"] .pubmed-mark { color: #8FB8E6; }
@media (prefers-color-scheme: dark) { :root:not([data-theme="light"]) .pubmed-mark { color: #8FB8E6; } }
.link-btn--pubmed:hover .pubmed-mark, .tag--pubmed .pubmed-mark { color: #20558A; }
.tag--pubmed { background: #fff; border-color: var(--azul-200); }
.tag--pubmed .pubmed-mark { font-size: .78rem; }

.footer-social { display: flex; gap: .5rem; margin-top: 1.25rem; }
.footer-social .link-btn { background: rgb(255 255 255 / .06); border-color: rgb(255 255 255 / .12); color: var(--azul-200); }
.social-row { display: flex; flex-wrap: wrap; gap: .5rem; }
.contact-card .social-row a.link-btn {
  display: inline-flex; align-items: center; gap: .5rem; padding: .55rem 1rem; border-radius: 999px;
  border: 1px solid var(--line); background: var(--bg-alt); text-decoration: none; color: var(--heading);
  transition: transform .25s var(--ease), background .2s, color .2s;
}
.contact-card .social-row a.link-btn:hover { color: #fff; }

/* Colaboradores en "Sobre nosotros" */
.about-collabs { margin-top: 1.75rem; padding-top: 1.5rem; border-top: 1px solid var(--line); }
.inst-list { display: flex; flex-wrap: wrap; gap: .5rem; margin: .9rem 0 1rem; }
.inst-chip { padding: .5rem .9rem; border-radius: 999px; background: var(--bg-alt); border: 1px solid var(--line); text-decoration: none; font: 600 .92rem/1.2 var(--font-display); color: var(--heading); transition: border-color .2s, transform .25s var(--ease); }
.inst-chip:hover { border-color: var(--genoma); transform: translateY(-2px); }
.inline-link { display: inline-flex; align-items: center; gap: .4rem; font-weight: 700; color: var(--accent-text); text-decoration: none; }
.inline-link svg { width: 1.05em; height: 1.05em; transition: transform .25s var(--ease); }
.inline-link:hover svg { transform: translateX(4px); }

/* Colaboradores: clínica e investigación unidas por el laboratorio (doble hebra en el centro) */
.collab-bridge { display: grid; grid-template-columns: minmax(0, 1fr) 128px minmax(0, 1fr); align-items: stretch; }
.collab-side { display: flex; flex-direction: column; padding: clamp(1.4rem, 3vw, 2rem); border-radius: var(--radius-lg); background: var(--surface); border: 1px solid var(--line); box-shadow: var(--shadow); }
.collab-side > .label { align-self: flex-start; }
.collab-side:only-child { grid-column: 1 / -1; }
.collab-place { margin-top: 1.4rem; padding-top: 1.4rem; border-top: 1px solid var(--line); }
.collab-place:last-child { flex: 1; display: flex; flex-direction: column; }
.collab-side > .label + .collab-place { margin-top: 1.25rem; padding-top: 0; border-top: 0; }
.collab-place h3 { margin: 0 0 .45rem; }
.collab-meta { display: block; margin-bottom: 1.1rem; font: 500 .75rem/1 var(--font-mono); letter-spacing: .14em; text-transform: uppercase; color: var(--text-soft); }
.collab-people { list-style: none; margin: 0; padding: 0; display: grid; grid-template-columns: repeat(auto-fill, minmax(min(100%, 200px), 1fr)); gap: .9rem 1.5rem; }
.collab-people strong { display: block; font: 600 1.02rem/1.25 var(--font-display); color: var(--heading); }
.collab-people span { font-size: .92rem; color: var(--text-soft); }
.collab-lines { display: flex; flex-wrap: wrap; align-items: center; gap: .6rem .75rem; margin-top: auto; padding-top: 1.4rem; }
.collab-lines-label { font: 500 .72rem/1 var(--font-mono); letter-spacing: .14em; text-transform: uppercase; color: var(--text-soft); }
.collab-line {
  display: inline-flex; align-items: center; gap: .55rem; padding: .3rem .9rem .3rem .3rem; border-radius: 999px;
  background: color-mix(in srgb, var(--c) 14%, transparent); color: var(--heading);
  font-size: .9rem; font-weight: 700; line-height: 1.25; text-decoration: none; transition: background .2s;
}
.collab-line:hover { background: color-mix(in srgb, var(--c) 24%, transparent); }
.collab-line b { flex: none; width: 1.75rem; height: 1.75rem; border-radius: 50%; display: grid; place-items: center; background: var(--c); color: var(--nucleo-deep); font: 600 .8rem/1 var(--font-mono); }
.collab-hub { position: relative; display: grid; place-items: center; background: linear-gradient(var(--genoma), var(--genoma)) center / 100% 2px no-repeat; }
.collab-hub::before, .collab-hub::after { content: ""; position: absolute; width: 3px; border-radius: 3px; background: var(--genoma); }
.collab-hub::before { top: 0; bottom: 0; left: calc(50% - 5px); }
.collab-hub::after { top: 0; bottom: 38%; left: calc(50% + 2px); background: var(--base-a); }
.collab-hub span {
  position: relative; z-index: 1; padding: .6rem .85rem; border-radius: 999px; background: var(--genoma); color: #fff;
  font: 600 .72rem/1 var(--font-mono); letter-spacing: .14em; text-transform: uppercase; white-space: nowrap;
  box-shadow: 0 0 0 6px var(--bg-alt);
}
@media (max-width: 860px) {
  .collab-bridge { grid-template-columns: minmax(0, 1fr); }
  .collab-hub { height: 76px; background-size: 2px 100%; }
  .collab-hub::before { top: calc(50% - 5px); bottom: auto; left: 0; right: 0; width: auto; height: 3px; }
  .collab-hub::after { top: calc(50% + 2px); bottom: auto; left: 0; right: 38%; width: auto; height: 3px; }
}

/* Financiación: logos oficiales siempre sobre blanco */
.funding-panel { margin: 0 auto; max-width: 980px; padding: clamp(20px, 4vw, 40px); border-radius: var(--radius-lg); background: #fff; border: 1px solid var(--line); box-shadow: var(--shadow); }
.funding-panel img { width: 100%; height: auto; }
.funding-panel figcaption { margin-top: 1.25rem; text-align: center; font-size: .85rem; line-height: 1.7; color: #46607F; }
.funding-panel figcaption a { color: #46607F; text-decoration: none; }
.funding-panel figcaption a:hover { color: var(--base-a-text); text-decoration: underline; }

/* Instagram */
.ig-grid { display: grid; grid-template-columns: repeat(auto-fill, minmax(min(100%, 320px), 1fr)); gap: clamp(16px, 2.5vw, 24px); align-items: start; }
.ig-slot { position: relative; aspect-ratio: 4 / 5; border-radius: var(--radius-lg); overflow: hidden; background: linear-gradient(135deg, var(--nucleo-deep), var(--genoma)); box-shadow: var(--shadow); }
.ig-slot .news-pattern { font-size: 1.05rem; }
.ig-badge { position: absolute; inset: 0; display: grid; place-items: center; color: #fff; }
.ig-badge svg { width: 64px; height: 64px; padding: 14px; border-radius: 22px; background: linear-gradient(45deg, #F58529, #DD2A7B 55%, #8134AF); box-shadow: 0 12px 30px rgb(0 0 0 / .3); transition: transform .5s var(--ease); }
.ig-slot:hover .ig-badge svg { transform: scale(1.08) rotate(-6deg); }
.ig-index { position: absolute; left: 14px; bottom: 12px; font-size: .75rem; letter-spacing: .1em; color: var(--azul-200); }
.ig-slot.is-embed { aspect-ratio: auto; background: transparent; box-shadow: none; overflow: visible; border-radius: 0; }
.ig-slot .instagram-media { margin: 0 !important; min-width: 0 !important; width: 100% !important; max-width: 100% !important; border-radius: var(--radius) !important; border-color: var(--line) !important; box-shadow: var(--shadow) !important; }
.ig-actions { display: flex; flex-wrap: wrap; gap: .75rem; align-items: center; margin-top: 1.75rem; }
.ig-actions .btn svg { width: 1.2em; height: 1.2em; }
.ig-actions .btn:hover svg { transform: none; }
.ig-consent { flex-basis: 100%; margin: 0; font-size: .85rem; color: var(--text-soft); }
#colaboradores .grid { align-items: start; }
.link-btn.is-pending { cursor: default; }
/* iconos alineados al pie de cada tarjeta aunque el cargo ocupe dos líneas */
.member:not(.member--pi) { display: flex; flex-direction: column; }
.member:not(.member--pi) .member-body { flex: 1; display: flex; flex-direction: column; }
.member:not(.member--pi) .member-links { margin-top: auto; padding-top: .85rem; }